Mamenchisaurus

A genus of sauropod dinosaurs from the Jurassic era of China
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This entry was edited and reviewed by the Institute of Vertebrate Paleontology and Paleoanthropology, Chinese Academy of Sciences, and verified by Science Encyclopedia China.
Mamenchisaurus is sauropods A genus of dinosaurs known for their very long necks, which make up almost half of their total body length. Several species have been assigned to Mamenxizaurus, and the attribution of individual species may be disputed. in Western China Its fossils have been found in several provinces. with Omeisaurus the kinship More recently, both belong to Mamensiidae. Most Mamenxisaurus can be over 20 meters long, and a few individuals can exceed 30 meters. Estimates of the weight of the species vary widely, with the smaller species thought to weigh less than 10 tons, while the largest species is thought to have probably weighed more than 60 tons.
Mamenxiosaur dinosaurs are not unique to East Asia. [13]
Chinese name
Mamenchisaurus
Foreign name
Mamenchisaurus
Type species
Construction of the Mamenxi dragon ( Mamenchisaurus constructus )
Sort position
Chordate - Vertebrate Subphyla - Sauroforma - Dipora - Dinoidea - Sauropyta - Sauropoda - Mamenxiidae

Mamenchisaurus

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Origin: Xinjiang, Gansu, Sichuan and Yunnan provinces of China.
Age: Late Jurassic Oxford Epoch Early Cretaceous Aptic age About 163 million to 113 million years ago.
Naming person and naming year: Yang Zhongjian 1954

Origin of name

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First Mamenchisaurus ( Mamenchisaurus The fossil was found on a highway construction site in Sichuan in 1952, and the incomplete skeleton was studied later and made famous in China in 1954 paleontologist Yang Zhongjian Named for construction Mamenxisaurus ( Mamenchisaurus constructus ). [14 ] The genus name "Mamenchi" means "Mamen Creek", where the fossil was found, and "saurus" means Latin " lizard And we generally translate reptiles as "dragons", so it is translated as "Mamenxi dragon". The fossils originally studied were located in Sichuan Province Yibin The construction site next to the Mamingxi ferry on the Jinsha River, but due to the accent of Mr. Yang Zhongjian, the "Mamingxi" was mistaken as "Mamenxi"; Species name construction ( constructus ) means that the first fossil was found in Construction site Unearthed. [1]

Morphological characteristics

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Neck minister (C-spine 17-19), Cervical rib It is relatively long, and the leading edge of the neck rib is dagger-shaped; The longest vertebra of the cervical spine is the single vertebra of the dorsal vertebra Average length 3.5-3.7 times; Both cervical and dorsal vertebrae are posterior concave; The sacral vertebra consists of 4-6 healed vertebrae, the nerve spines of the sacral vertebrae are healed together. [2]

Species discovery

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Construction of the Mamenxi dragon

Mamenchisaurus constructus was first found in China in 1952 Sichuan It was found at a construction site on the Itang Highway in the province. The fossil site belongs to the Shangshaximiao Formation and dates back at least to the Late Jurassic. In 1954, Yang Zhongjian, a famous Chinese paleontologist, studied this part of the skeleton and named it Mamenxiosaur.
holotype (IVPP V 790) mutilated and disorganized. Materials included: 5 dorsal vertebrae, 30 Coccygeal vertebra , rib fragments, dorsal nerve spines and Pulse arc . There were 14 cervical vertebrae, but none of them were complete, and Young noted that some cervical vertebrae may have been missing and estimated the entire neck to be 4.67 meters. There are also partial limbs, including: two femurs, a complete tibia, fibula , talus , Metatarsal bone , phalanx And claws. The skull, forelimbs, and waistband were not preserved. It is estimated that the length of the Mamenxisaurus was about 13 to 15 meters and the mass was 5 tons. [1-2]
Construction of the Mamenxi dragon

Mamenxizaurus hechuanensis

In 1972, a second species of Mamenxiosaurus was introduced by Yang Zhongjian He Zhao Xijin It was named Mamenchisaurus hochuanensis. These fossils are in China Chongqing The city Hechuan District Found near a village. The fossil site of Mamenxiosaur in Hechuan also belongs to Shangshaximiao Formation, which is very close to the horizon where the holotype specimens of Mamenxiosaur were constructed, and can be traced back to at least the Late Jurassic. Locals first discovered the remains sometime before the founding of New China, however, the remains were eventually left in place and excavation work did not begin until 1957.
The holotype (CCG V 20401) consists of an almost complete associated spine, consisting of 19 almost completely preserved elongated cervical vertebrae, 12 dorsal vertebrae, 4 sacral vertebrae, and 35 caudal vertebrae, with only the last few caudal vertebrae missing. The skeleton is also missing most of the forelimbs and skull. The total length of Mamenxiosaur was about 21 to 22 meters, with a neck length of 9.3 meters. When the Hechuan Mamenxiosaur was first described, it was the largest known in China sauropods Animals. Yang and Zhao estimate that Mamenxiosaur Hechuan weighed 45 tons. However, later weight Estimated value Lower, 14-18.2 tons. [2-3]
In August 2022, Asia integrity Dinosaur fossil The largest of the holotype specimens, the Hechuan Mamenxiosaur, completed the first overhaul since the fossil installation in 1965. The specimen of Mamenxisaurus hechuanensis preserved 35 tail vertebrae, and the tail vertebrae were restored by 55 sections. After restoration, the skeleton length was "longer" by 2 meters, and the total length was increased to 24 meters Chengdu Museum of Natural History (Chengdu University of Technology Museum) Second floor "Dragon Travel Sichuan-Chongqing" exhibition hall. [12]
Mamenxizaurus hechuanensis

The Gamamentesaurus

In 1993, the third species of Mamenxisaurus -- The Gamamentesaurus (Mamenchisaurussinocanadorum) is described. The species name refers to Chinese and Chinese Canada the paleontology Joint study project. The remains were discovered in 1987, when a neck rib was seen protruding from a cliff. The fossil site is located in Xinjiang Junggar Basin General Temple area on the east margin; From the upper Shishugou Formation (161.2 Ma to 158.7 Ma) of the early Late Jurassic. [4] The specimen (IVPP V 10603) consisted of a well-preserved joint sequence of the first four cervical vertebrae Lower jaw Composed of bone and other skull material. mandible It is 60.3 cm long and has 19 teeth. with diplodocus In contrast to the more square mandible of the class, the anterior mandible of this specimen is presented Bevel Angle . vertebra The neural arch and vertebral body healed well, indicating that the animal was mature. In the description Sauropod dinosaurs The species has the longest neck rib, 4.1 m long. The average length of the cervical vertebrae of Gabamenxisaurus was 1.19 times that of that of Hechuan Mamenxisaurus. Based on this, the researchers estimated the full length of the specimen to be 26 meters. It is estimated that this dinosaur had a neck of 12 Meter length . [2] [5]
The Gamamentesaurus

Mamensaurus yangi

The Shangshaximiao Formation was described in 1996 Mamensaurus yangi (Mamenchisaurus youngi). His remains were found in 1987 in Sichuan Zigong Xinmin Township a village near one Stone pit Found. The species was named in honor of Mr. Yang Zhongjian.
The holotype specimen (ZDM 0083) is very complete and mostly associated, preserved from the head to the 8th Caudal vertebra All vertebral bodies of; This species has 18 cervical vertebrae. And also saved it Shoulder girdle , waistband, and material for all limbs. The specimen also preserves a nearly complete skull. premaxilla It has four teeth, Upper jaw bone It has 18 teeth, Dental bone It has 22-24 teeth. With a length of 16 meters and a neck length of 6.5 meters, it is one of the smaller species of Mamenxiosaur. [2] [6]
Mamensaurus yangi

Mamenxisaurus anyue

In 1996, He Xinlu et al. described Mamenxisaurus anyue (Mamenchisaurusanyuensis). His remains were found in 1987 Sichuan basin Anyue county Longqiao township Two nearby locations were found. In one quarry, at least 5-6 individuals were found. In the second quarry, four individuals were found. One of the specimens, AL001, was a holotype, preserving two-thirds of the associated bones of the whole body. Other specimens, AL002, AL003, and AL101-106, were also reported, which provided additional skeletal information. The name of the species is related to Anyue County, where the species was found. Mamenxisaurus anyuensis was found at the top of Suining Formation and at the bottom of Penglaizhen Formation. Uranium lead The age of Mamenxisaurus Anyue from Suining Formation is 114.4 Ma. Since this would make it about 30 million years younger than the other Mamenxiosaur species, there is some doubt whether Anyue Mamenxiosaur is a member of the genus Mamenxiosaur.
The holotype specimens preserved 8 posterior cervical vertebrae, 12 dorsal vertebrae, 5 sacral vertebrae, several caudal vertebrae, and a complete girdle. Another specimen (AL102) retained five associated cervical vertebrae in the middle cervical vertebrae. In addition to the phalanx, other parts of the forelimb were found. This species has many morphological similarities with Mamenxiosaurus hechuanensis. The length of this species is estimated to be 21 meters. [2] [7]

Jingyan Mamenxisaurus

In 1998, Zhang Yihong, Li Kui and Zeng Qinghua described Jingyan County Mamenchisaurus jingyanensis. The holotype specimen (CV00734) was found Jingyan county The village of Meiwang. About 10 kilometers from Maywan Sanjiang Town Pengjiazhai Another paratype (JV002) was found in the Western Mountains. The third specimen (CV00219) was found near Dujia village, about 10 km from Sanjiang. The fossils are located in the Upper Shaximiao Formation in the Sichuan Basin. The species name refers to the fact that most of the specimens were excavated from Jingyan County.
The holotype specimen consists of parts scapula And complete coracoid , forelimb, complete ischium , relatively intact skull and hyoid bone. Paratypes include three anterior cervical vertebrae, several weathered dorsal vertebrae, various caudal vertebrae, relatively intact hind and forelimbs, and isolated teeth. The third specimen preserves a cervical vertebra with associated cervical ribs, four fused sacral vertebrae, several coccygeal vertebrae, a complete scapula, various limb bones, and isolated teeth. The length of the skull is 55 cm. The prejaw has four teeth, the maxilla has 14-16 teeth, and the dental bone has 17-19 teeth. The length of this species is estimated to be between 20 and 26 meters. [2] [8]
Jingyan Mamenxisaurus

Other discoveries of Mamensaurus

Over the years, other species of Mamenxisaurus have also been named. Some of these species are based on fragmentary remains that are considered invalid. In some cases, species from other genera have also been migrated to Mamenchiosaurus, but on migrations effectiveness There are different opinions.
In 1976, researchers named the four specimens found in the Shangshaximiao Formation as Fuxi zigongosaurus . [9] Since then, other researchers have disagreed about whether the genus is valid. A 1983 study relocated the remains to Omeisaurus Belong to the year. In a 1996 study, the remains were transferred to the genus Mamenxisaurus, which was named Mamenxisaurus Fuxiensis. [10]
In 1998, Zhang Yihong, Li Kui, and Zeng Qinghua mentioned it briefly in their descriptions of Jingyan Mamenxisaurus Mamenxisaurus guangyuanensis No details were given. Since then, other research views have differed on the validity of this species. [8]
In 2004, a study by Fang Xiaosi et al. was published in the journal Yunnan (Province) Mamenxisaurus yunnanensis unearthed from Anning Formation, province. [11]
The ancient biota
The fossils of Mamenxiosaur have been found in Shangshaximiao Formation, Shishugou Formation, Suining Formation, Penglaizhen Formation and Anning Formation, dating from Late Jurassic to Early Cretaceous. In different strata, the assemblage of fossils is different, reflecting different paleontological communities. With the late Jurassic found in Sichuan Mamenxiosaur fossil It was excavated from the same level Dinosaur Includes: sauropods Emei Dragon, Mamenxi dragon, theropoda the Yongchuanosaurus , Sinraptor , ornithischia the Yandusaurus , Chungkingosaurus , Tuojiangosaurus Etc., and a few more testudines and crocodilians Of the fossil.
Also unearthed from the Late Jurassic Shishugou Formation in Xinjiang are: Sauropod Mamenxisaurus, bellusaurus Theropod corythosaurus , Janchosaurus , Sinraptor, haplosteophosaurus bird-hip Colorface Ceratops , Hidden dragon , and several pterosaur And crocodiles.
Mamenxisaurus Anyue is the only one in Mamenxisaurus unearthed in the Early Cretaceous stratum kinship adjacent Qijianglong .
The size ratio of different species of Mamenchisaurus